Up + Down - Heal. Health can be restored up to the top of the current bar, but the Life Counter cannot be turned by healing. Taking damage interrupts healing.
Left + Right - Taunt. Taunting reluctant enemies aggravates them and makes them more likely to attack, but the player is vulnerable whilst taunting.
A - A light attack that deals little damage but is very fast.
A + Up - A heavy attack that is very slow and easily interrupted, but does large amounts of damage.
A + Opposite direction to enemy - Fire a Thornax fruit. The blue Thornax meter must be completely full to perform this move, and recharges whilst not in use.
S - A blocking maneuver that can parry enemy blows.
S + Up or Down - Flip/cartwheel sideways and back in the chosen direction, reducing the chance of being struck with an attack.
S + Opposite direction to enemy - Back-flip away from the enemy and automatically fire a Thornax upon landing, performing Gresh's signature move. The blue Thornax meter must be completely full to perform this move.